... I think it's more important a lens is being used for its intended purpose once in a while. ...
I agree with those who say use the lens and if that isn't happening then with those who say to exercise the lens. I think Bjørn mentioned lenses getting stiff above from lack of use. This is probably fairly common. If the lube isn't normal or stiff it may very slowly flow. Using normally or exercising may redistribute the lubricant and keep it from getting where it's not supposed to be like the aperture blades.
Excessive heat hasn't been mentioned. Most know not to keep a camera or lenses in the trunk or cabin of a car or truck but when on vacation or trying to save money by not using air conditioning heat may affect lubricants, flow and cause lube contamination of the aperture blades. Much of Southern California is hot and dry much of the year with cool moist winters. Houses with forced heat and air conditioning keeps the temperatures moderate and humidity low.
Foam cases: there are different kinds of foam. They can look the same, e.g. charcoal gray. Some can off gas and fog lenses. I remember no specifics materials used. Cheep cases and unknown sources may result in using the wrong foam and damage.
My 2 cents is...
1) Store most lenses face down with caps. Dividers that hold lenses in a drawer can help.
2) Telephotos if they can't be stored face down turn regularly.
3) Avoid high humidity (not usually a problem for me, inland So. California).
4) Avoid high temperatures. Maybe set air conditioning at 78*F (25.5*C) and pay the electric bill rather than lens repair. Cool only one area if heat and air is installed for this option.
5) Use approved foam in cases. Avoid any storage that may off gas.
I'm sure there are more conditions to use or avoid. Got to run.
Dave Hartman