If you trip the aperture lever with a finger, several times in a row, to simulate when it is actually mounted on the camera and stopping down and opening again, you will most likely see that it makes a circular opening,,,
Several reasons for the issue with it not looking round all the time:
The return spring is not very strong on these 125mm APO Lanthars,,,
The activating mechanism for the aperture has to go deep inside the lens due to the very long extension to reach 1:1
The movement is reversed inside mount, it's a universal lens build to fit many camera systems, unfortunately not with Nikon F as the main aperture stop down rotation direction,,,
The reversal movement is a strange rotating ring activated reversed by a rotating plate and a hook-shaped thing:
_EGL2051 by
Erik Gunst Lund, on Flickr
Here is an image of the return spring for the aperture movement, here it is relocated due to it has one of Bjørns CPU build in (Ai-P)
_EGL2050 by
Erik Gunst Lund, on Flickr
Here you can see the CPU Contact block
_EGL2057 by
Erik Gunst Lund, on Flickr
Just enjoy the lens