Looked through the portfolio of that session and all three factors contribute, but the influence of dead branches probably was instrumental.
Here is another, from the setting-up stage;
At the beginning I worked mostly in the visible domain as the slowly setting sun imparted a magical spell of reds into the dense forest. When the sun rays faded, IR became the better option. My camera at the time, the D2H, allowed both techniques just by replacing filters in the drawer of the 300 Nikkor I used as my tool of choice for the day.